To the taxonomy of the subfamily Aradinae (Heteroptera, Aradidae)
Aradinae (Hemiptera, Heteroptera, Pentatomomorpha, Aradoidea, Aradidae – “flat bugs”) is mostly Holarctic (temperate and subtropics of Eurasia and North America) group. Here I have tried to summarize information about the taxonomy of the subfamily.
The most important catalogs are:
Berend Aukema (ed.). Catalogue of Palaearctic Heteroptera. 1995-2013 (the electronic database is supplemented to date). Available online:
https://catpalhet.linnaeus.naturalis.nl/linnaeus_ng/app/views/introduction/topic.php?id=9
Thomas J. Henry, Richard C. Froeschner (eds.). 1988 (eBook Published 17 July 2019). Catalog of the Heteroptera or True Bugs, of Canada and the Continental United States. DOI: https://doi.org/10.1201/9781351070447
María del Carmen Coscarón, Eugenia Fernanda Contreras. 2012. Catalog of Aradidae (Hemiptera: Heteroptera) for the Neotropical Region. DOI: https://doi.org/10.11646/zootaxa.3466.1.1
J. G. Pendergrast. 1968. The Aradidae of New Zealand (Hemiptera, Heteroptera) III – The Aradinae and Calisiinae. Transactions of the Royal Society of New Zealand: Zoology, Volume 10, Issue 10. Available online:
https://paperspast.natlib.govt.nz/periodicals/TRSZOO19680122.2.2
Aradinae – Atlas of Living Australia.
Some identification guides (continental or country level, most of them somewhat outdated):
Heiss, E. & J. Péricart. 2007. Hemiptères Aradidae Piesmatidae et Dipsocoromorphes euro-méditerranéens. Faune de France n° 91. ISBN: 978-2-9030-5229-4 (Western Palearctic)
Ryuichi Matsuda. 1977. The Insects and Arachnids of Canada, part 3. The Aradidae of Canada. Available online: https://publications.gc.ca/site/eng/9.811337/publication.html (Canada
and adjacent states of the US)
J. R. de la Torre-Bueno. 1939. A Synopsis of the Hemiptera Heteroptera of America North of Mexico – Part I – Families Scutelleridae, Cydnidae, Pentatomidae, Aradidae, Dysodiiclae and Termitaphididae. Available online:
https://www.biodiversitylibrary.org/item/205520#page/276/mode/1up (the US and Canada)
Hsiao, T.Y., S.Z. Ren, L.Y. Zheng, H.L. Jing, H.G. Zou & S.L. Liu. 1981. A handbook for the determination of the Chinese Hemiptera-Heteroptera. Volume II: i-iv, 1-654 (China)
Y Nakatani, K Kohno, T Ishikawa, Mikio Takai, Tomohide Yasunaga, Ernst Heiss. 2012. Terrestrial Heteropterans: A Field Guide to Japanese Bugs, Volume 3. ISBN: 9784881371688 (Japan)
Wonwoong Kim, Minsuk Oh, WonGun Kim, Seunghwan Lee. 2023. New records of genus Aradus Fabricius (Hemiptera: Aradidae) from Korea, with first male description of Aradus gretae Kiritshenko. Journal of Asia-Pacific Biodiversity, Volume 16, Issue 2. Pages 157-162. ISSN 2287-884X, https://doi.org/10.1016/j.japb.2022.12.004. (https://www.sciencedirect.com/science/article/pii/S2287884X22001352) (Korea)
In the following preliminary list of species of Aradinae of the World fauna I may have missed some tropical species. The status of some taxa remains debatable. The list is available as a table (https://docs.google.com/spreadsheets/d/1QdLbUQZ9rm_jaDVs4YACV3_PD7gDrCujg5U8JEl2CUw/edit?usp=sharing). Unfortunately, only some Palaearctic species may be referred to “species groups” (according to Heiss & Péricart, 2007).
SUBFAMILY ARADINAE
Genus Aradiolus Kormilev 1967 (Neotropic)
- chemsaki Kormilev, 1980
- paradoxus Kormilev, 1967
Genus Aradus Fabricius, 1803 (World)
Subgenus Aradus s.str. (World)
Species known from Western Palaearctic
betulae group
- betulae (Linnaeus, 1758)
- brenskei Reuter, 1884
- caucasicus Kolenati, 1857
- elburzanus Heiss, 2004
- hieroglyphicus J. Sahlberg, 1878
- krueperi Reuter, 1884
- maghrebinus Heiss, 2003
- obscurus Vásárhelyi, 1988
- ribauti Wagner, 1956
- seidenstueckeri Heiss, 1989
pictus group
- annulicornis Fabricius, 1803
- horvathi Vásárhelyi, 1984
- obtectus Vásárhelyi, 1988
- pictus Baerensprung, 1859
corticalis group
- betulinus Fallén, 1807
- brevicollis Fallén, 1807
- corticalis (Linnaeus, 1758)
conspicuus group
- cilicicus Heiss, 1990
- conspicuus Herrich-Schaeffer, 1835
- inopinus Kiritshenko, 1955
erosus group
- erosus Fallén, 1807
- montandoni Reuter, 1885
lugubris group
- angularis J. Sahlberg, 1886
- aterrimus Fieber, 1864
- bimaculatus Reuter, 1872
- canariensis Kormilev, 1954
- laeviusculus Reuter, 1875
- lugubris Fallén, 1807
depressus group
- depressus (Fabricius, 1794) (including depressus depressus (Fabricius, 1794) and depressus leptocerus Horváth, 1882)
- versicolor Herrich-Schaeffer, 1835
- serbicus Horváth, 1888
- somcheticus Kiritshenko, 1913
- truncatus Fieber, 1860
mirus group
- mirus Bergroth, 1894
- reuterianus Puton, 1875
stenopterus group
- cinnamomeus Panzer, 1806
- graecus Heiss, 1997
- lauri Noualhier, 1893
- safavii Hoberlandt, 1974
- stenopterus Bergroth, 1887
- tauricus Jakovlev, 1907
species incertae sedis
- crenaticollis R.F. Sahlberg, 1848
- distinctus Fieber, 1860
- flavicornis Dalman, 1823
- kuthyi Horváth, 1899
- pallescens Herrich-Schaeffer, 1840 (including pallescens pallescens Herrich-Schaeffer, 1840 and pallescens frigidus Kiritshenko, 1913)
- rysakovi Kiritshenko, 1951
- signaticornis R.F. Sahlberg, 1848
Species known from Eastern Palaearctic only
- bergrothianus Kiritshenko, 1913
- candidatus Bergroth, 1889
- compar Kiritshenko, 1913
- consentaneus Horváth, 1905
- czerskii Kiritshenko, 1915
- dembickyi Heiss, 2007
- discompar Hsiao, 1964
- dissimilis A. Costa, 1847
- dissors Kiritshenko, 1913
- emeiensis Heiss, 2008
- esakii Kormilev & Heiss, 1976
- formosanus Heiss, 2009
- gaoligongensis Bai, Heiss & Cai, 2010
- gretae Kiritshenko, 1955
- herculeanus Kiritshenko, 1913
- holzschuhi Heiss, 2003
- honshuensis Heiss & Shono, 2005
- komarowii Jakovlev, 1885
- kyrgyzstanus Heiss, 2004
- miyamotoi Heiss & Shono, 2003
- nipponicus Kormilev, 1955
- omeiensis Hsiao, 1964
- orientalis Bergroth, 1885
- oroszi Vásárhelyi, 2016
- parayunnanus Heiss, 2010
- pictellus Kerzhner, 1973
- poppiusi Kiritshenko, 1913
- pulchellus J. Sahlberg, 1878
- quinlingshanensis Heiss, 2003
- semilacer Kiritshenko, 1913
- setiger Kiritshenko, 1913
- sinensis Kormilev, 1955
- smetanai Heiss, 2003
- spinicollis Jakovlev, 1880
- tonkinensis Kormilev & Heiss, 1976
- transiens Kiritshenko, 1913
- trapezonotus Heiss & Shono, 2005
- turnai Heiss, 2008
- unicolor Kiritshenko, 1913
- ussurensis Jakovlev, 1876
- vitiosus Jakovlev, 1889
- yunnanus Heiss, 2003
- zhengi Heiss, 2001
Species known from South Asia only (excl. candidatus (also Eastern Palaearctic))
- abnormis Bergroth, 1889
- burmensis Kormilev, 1976
- malaisei Kormilev, 1976
Species known from Nearctic (the US and Canada) (excl. Holarctic lugubris)
- abbas Bergroth, 1889
- acutus Say, 1832
- aequalis Say, 1832
- alaskanus Kormilev and Heiss, 1979
- ampliatus Uhler, 1876
- antennalis Parshley, 1921
- apicalis Van Duzee, 1920
- approximatus Parshley, 1921
- arizonicus Parshley, 1921
- barberi Kormilev, 1966
- basalis Parshley, 1921
- behrensi Bergroth, 1886
- blaisdelli Van Duzee, 1920
- borealis Heidemann, 1909
- breviatus Bergroth, 1887
- brevicornis Kormilev, 1980
- brunnicornis Blatchley, 1926
- carolinensis Kormilev, 1964
- cincticornis Bergroth, 1906
- coarctatus Heidemann, 1907
- coloradensis Kormilev, 1964
- compressus Heidemann, 1907
- concinnus Bergroth, 1892
- consors Parshley, 1921
- crenatus Say, 1832
- curticollis Bergroth, 1913
- debilis Uhler, 1876
- depictus Van Duzee, 1917
- duzeei Bergroth, 1892
- evermanni Van Duzee, 1920
- falleni Stål, 1860
- funestus Bergroth, 1913
- urnissi Usinger, 1936
- furvus Parshley, 1921
- fuscipennis Usinger, 1936
- fuscomaculatus Stål, 1859
- gracilicornis Stål, 1873
- gracilis Parshley, 1929
- hesperius Parshley, 1921
- implanus Parshley, 1921
- inornatus Uhler, 1876
- insignitus Parshley, 1921
- insolitus Van Duzee, 1916
- intectus Parshley, 1921
- intermedius Usinger, 1936
- kormilevi Heiss, 1980
- lawrencei Kormilev, 1966
- leachi Van Duzee, 1929
- linsleyi Usinger, 1936
- marginatus Uhler, 1893
- medioximus Parshley, 1921
- montanus Bergroth, 1913
- occidentalis Kormilev, 1980
- opertaneus Parshley, 1921
- orbiculus Van Duzee, 1920
- oregonicus Kormilev, 1978
- ornatus Say, 1832
- ovatus Kormilev, 1966
- oviventris Kormilev, 1966
- paganicus Parshley, 1929
- pannosus Van Duzee, 1920 (including pannosus pannosus Van Duzee, 1920 and pannosus incomptus Parshley, 1921)
- parshleyi Van Duzee, 1920
- parvicornis Parshley, 1921
- patibulus Van Duzee, 1927
- persimilis Van Duzee, 1916
- proboscideus Walker, 1873
- quadrilineatus Say, 1825
- robustus Uhler, 1871 (including robustus robustus Uhler, 1871 and robustus insignis Parshley, 1921)
- saileri Kormilev, 1966
- saskatchewanensis Matsuda, 1980
- serratus Usinger, 1936
- shermani Heidemann, 1907
- signaticornis Sahlberg, 1848
- similis Say, 1832
- snowi Van Duzee, 1920
- subruficeps Hussey, 1953
- taylori Van Duzee, 1920
- tuberculifer Kirby, 1837
- uniannulatus Parshley, 1921
- uniformis Heidemann, 1904
- vadosus Van Duzee, 1920
- vandykei Van Duzee, 1927
Species known from Neotropics only (excl. falleni and gracilicornis that are also occur in the US)
- angustellus (Blanchard, 1852)
- brasiliensis Usinger, 1940
- compressicornis Stål, 1873
- fronterana Drake, 1942
- granuliger Kormilev, 1980
- mexicanus Usinger, 1936
- pallidicornis Stål, 1873
- penningtoni Drake, 1942
- pericarti Heiss, 1994
- testaceus Kormilev, 1980
Species known from Australia and New Zealand
- albicornis (Walker, 1873)
- australis Erichson, 1842
- erraticus Kormilev, 1966
- fuscicornis Kormilev, 1966
- leucotelus (Walker, 1873)
Species known from Sub-Saharan Africa only
- gracilipes Linnavuori, 1978
- zavattarii Mancini,1939
Subgenus Lobataradus Heiss, 2008 (Estern Palaearctic, South Asia)
- bilobatus Heiss & Shono, 2005
- khuntanus Heiss, 2015
species inquirendae (type specimens are lost, maybe not Aradidae)
- griseus (Fabricius, 1794) (Acanthia grisea Fabricius, 1794)
- nigricornis (Fabricius, 1794) (Acanthia nigricornis Fabricius, 1794)
- tridentatus (Geoffroy, 1785) (Cimex tridentatus Geoffroy in Fourcroy, 1785)
Genus Miraradus Vásárhelyi, 1980 (Estern Palaearctic, South Asia)
- angulatus Heiss, 2009
- foliaceus (Kormilev, 1957)
- mirabilis (Bergroth, 1892)
- oervendetes Vásárhelyi, 1980
Genus Quilnus Stål, 1873 (Holarctic)
Species known from Western Palaearctic
- cedri (Puton, 1873)
- discedens (Horváth, 1911)
- marcosi Heiss & Baena, 2006
- mediterraneus Heiss, 1989
- parvicollis (Stål, 1873)
- subsimilis (Horváth, 1911)
Species known from Estern Palaearctic
- brevirostris (Horváth, 1901)
Species known from Nearctic
- heidemanni Bergroth, 1906
- niger Stål, 1873
- nigrinus Parshley, 1921 (including nigrinus nigrinus Parshley, 1921 and nigrinus canadensis Parshley, 1929)
- usingeri Kormilev, 1978